A pack of wolves have been put down at a Scots wildlife park.
The animals were euthanised following "complications" during an operation on the alpha male wolf, Loki, at Camperdown Wildlife Centre in Dundee. Despite Loki's initially successful operation, there were "subsequent complications" - and park bosses took the "difficult decision" to save the animal "from further pain" on Monday.
However, following the procedure, the other four wolves in the pack were showing "anxious and abnormal behaviours" - and so the animals were also then put to sleep. Bosses at Camperdown Wildlife Centre say the course of action was a "last resort" and that the team are "absolutely heartbroken".