Ben O'Connor has retained the leader's red, and Kaden Groves the green of the best sprinter, but Australia's hold on the three jerseys of the Vuelta a Espana is over after one day with Jay Vine dethroned as King of the Mountains.
Vine has been overtaken by his UAE Team Emirates teammate Marc Soler, who now leads by a point in the race for the polka dot jersey after he got into the main breakaway in the 18th stage, but Vine did not.
The combative Spaniard took full advantage, claiming all 15 points on offer in a 179.5km mid-mountain stage through Basque Country roads won by local rider Urko Berrade with a late solo break.