Doom: The Dark Ages' fifth mission introduces you to your dragon, letting you duke it out in the air, but also land at specific locations unlike the Atlan missions. This makes finding the Holy City of Aratum secrets and collectibles much more complicated, as you're no longer dealing with linear corridors.
After searching high and low, I've found all the Doom: The Dark Ages secrets in The Holy City of Aratum mission, including the secret dragon landing zones. Below you'll find all the secrets and collectibles laid out in chronological order, though keep in mind that this mission lets you tackle its main objectives in roughly any order, so I'll explain how that changes things.
