Canada zoo finds escaped wolf pups in moment of joy tinged with tragedy

Wolf pups in Minnesota. Two pups from the Greater Vancouver Zoo were found after four days on the loose. Photograph: All Canada Photos/Alamy

Emotions are bittersweet at a Canadian zoo after a runaway wolf pup was safely located after four days on the loose, but another was found dead along a road.

Conservation officers and zoo staff in Canada have spent the last four days searching for a runaway wolf after mysterious break-in freed a pack of the predators from the popular zoo.