Boris Johnson has suffered a brutal double defeat at the polls as voters turned their back on the Tories in two critical by-elections.
In a night of high drama, Labour won back the Red Wall seat of Wakefield and the Liberal Democrats overturned a 24,000 Tory majority in Tiverton and Honiton.
Conservative chairman Oliver Dowden quit this morning with a warning: "We cannot carry on with business as usual."