John Swinney has said that relations between the Scottish and UK Governments were "pretty poor" when the pandemic hit because of Brexit.
The former SNP Deputy First Minister told the UK Covid Inquiry in London that relationships between Westminster and Holyrood had been harmed by the prospect of Britain leaving the EU with no deal.
Swinney was speaking at the inquiry directly after former First Minister Nicola Sturgeon had answered questions. Ex-Scottish Health Secretary Jeane Freeman had a hearing on Wednesday.